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Dmanisi (tr, ˈdmɑnisi, Başkeçid) is a town and archaeological site in the Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. The hominin site is dated to 1.8 million years ago. It was the earliest known evidence of hominins outside Africa before stone tools dated to 2.1 million years were discovered in 2018 in Shangchen, China. A series of skulls which had diverse physical traits, discovered at Dmanisi in the early 2010s, led to the hypothesis that many separate species in the genus Homo were in fact a single lineage. Also known as Skull 5, D4500 is the fifth skull to be discovered in Dmanisi. The area around the town of Dmanisi has been settled since the Early Bronze Age. In the 6th century an Orthodox Christian cathedral named "Dmanisi Sioni" was built there. The oldest written record of the town is in the 9th century as a possession of the Arab emirate of Tbilisi. Located on the confluence of trading routes and cultural influences, Dmanisi was particularly important, growing into a major commercial center of medieval Georgia. The town was taken by the Seljuk Turks in the 1080s and by the Georgian kings David the Builder and Demetrios I between 1123 and 1125. The Turco-Mongol armies under Timur laid waste to the town in the 14th century. Sacked again by the Turkomans in 1486, Dmanisi never recovered and declined to a scarcely inhabited village by the 18th century. The castle was controlled by the House of Orbeliani. Dmanisi historic site Extensive archaeological studies began in the area in 1936 and continued in the 1960s. Beyond a rich collection of ancient and medieval artifacts and the ruins of various buildings and structures, unique remains of prehistoric animals and humans have been unearthed. Some of the animal bones were identified by the Georgian paleontologist A. Vekua with the teeth of the extinct rhino Dicerorhinus etruscus etruscus in 1983. This species dates back presumably to the early Pleistocene epoch.

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Mitochondrial Pseudogenes Suggest Repeated Inter-Species Hybridization among Direct Human Ancestors

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The hypothesis that the evolution of humans involves hybridization between diverged species has been actively debated in recent years. We present the following novel evidence in support of this hypothesis: the analysis of nuclear pseudogenes of mtDNA ("NUM ...

MDPI2022

Early human migrations

Early human migrations are the earliest migrations and expansions of archaic and modern humans across continents. They are believed to have begun approximately 2 million years ago with the early expansions out of Africa by Homo erectus. This initial migration was followed by other archaic humans including H. heidelbergensis, which lived around 500,000 years ago and was the likely ancestor of Denisovans and Neanderthals as well as modern humans. Early hominids had likely crossed land bridges that have now sunk.

Homo erectus

Homo erectus (ˌhoʊmoʊ_ə'rɛktəs; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Its specimens are among the first recognizable members of the genus Homo. Several human species, such as H. heidelbergensis and H. antecessor, appear to have evolved from H. erectus, and Neanderthals, Denisovans, and modern humans are in turn generally considered to have evolved from H. heidelbergensis. H.

Dmanisi hominins

The Dmanisi hominins, Dmanisi people, or Dmanisi man were a population of Early Pleistocene hominins whose fossils have been recovered at Dmanisi, Georgia. The fossils and stone tools recovered at Dmanisi range in age from 1.85 to 1.77 million years old, making the Dmanisi hominins the earliest well-dated hominin fossils in Eurasia and the best preserved fossils of early Homo from a single site so early in time, though earlier fossils and artifacts have been found in Asia.