Corticobulbar tract

In neuroanatomy, the corticobulbar (or corticonuclear) tract is a two-neuron white matter motor pathway connecting the motor cortex in the cerebral cortex to the medullary pyramids, which are part of the brainstem's medulla oblongata (also called "bulbar") region, and are primarily involved in carrying the motor function of the non-oculomotor cranial nerves. The corticobulbar tract is one of the pyramidal tracts, the other being the corticospinal tract. The corticobulbar tract originates in the primary motor cortex of the frontal lobe, just superior to the lateral fissure and rostral to the central sulcus in the precentral gyrus (Brodmann area 4). The corticobulbar tract however also includes fibres from disparate regions from across the cerebral cortex (not limited to the frontal lobes). The tract descends through the corona radiata and then the genu of the internal capsule (with a few fibers in the posterior limb of the internal capsule) to the midbrain. In the midbrain, the internal capsule becomes the cerebral peduncles. The white matter is located in the ventral portion of the cerebral peduncles, called the crus cerebri. The middle third of the crus cerebri contains the corticobulbar and corticospinal fibers. The corticobulbar fibers exit at the appropriate level of the brainstem to synapse on the lower motor neurons of the cranial nerves. In addition to endings in these motor neurons, fibers of the corticobulbar tract also end in the sensory nuclei of the brainstem including gracile nucleus, cuneate nucleus, solitary nucleus, and all trigeminal nuclei. Only 50% of the corticobulbar fibers decussate, in contrast to those of the corticospinal tract where most decussate: cranial nerve nuclei innervating skeletal muscle thereby generally receive bilateral first-order neuron innervation (i.e. from both hemispheres). The corticobulbar tract is composed of the upper motor neurons of the cranial nerves. The muscles of the face, head and neck are controlled by the corticobulbar system, which terminates on motor neurons within brainstem motor nuclei.