Concept

Gavialosuchus

Summary
Gavialosuchus is an extinct genus of gavialoid crocodylian from the early Miocene of Europe. Currently only one species is recognized, as a few other species of Gavialosuchus have since been reclassified to other genera. The type species, G. eggenburgensis, is known from the early Miocene of Austria. Two other species - G. americanus and G. carolinensis - have since been reclassified to other genera. Myrick Jr. (2001) proposed synonymizing Gavialosuchus americanus with Thecachampsa antiqua. Piras et al. (2007) advocated transferring both G. americanus and G. carolinensis to Thecachampsa as distinct species of the latter genus, however. Jouve et al. (2008) retained G. americanus in Gavialosuchus and found it to be the sister group of G. eggenburgensis (G. carolinensis was not discussed). However, Jouve et al. (2008) didn't test Thecachampsa antiqua in their phylogenetic analysis. Shan et al. (2009) found that G. americanus and G. eggenburgensis are not sister taxa. However, they didn't include T. antiqua and G. carolinensis in their analysis. Christopher A. Brochu and Glenn W. Storrs (2012) tested all four species, along with other crocodyloids, and found relatively strong support for Piras et al. (2007) suggestion. Weems (2018) agreed with Piras et al. (2007) and Brochu & Storrs (2012) at americanus and carolinensis are part of Thecachampsa. Below is a cladogram based morphological studies comparing skeletal features that shows Gavialosuchus as a member of Tomistominae, related to the false gharial: Based on morphological studies of extinct taxa, the tomistomines (including the living false gharial) were long thought to be classified as crocodiles and not closely related to gavialoids. However, recent molecular studies using DNA sequencing have consistently indicated that the false gharial (Tomistoma) (and by inference other related extinct forms in Tomistominae) actually belong to Gavialoidea.
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