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The order Pinales in the division Pinophyta, class Pinopsida, comprises all the extant conifers. The distinguishing characteristic is the reproductive structure known as a cone produced by all Pinales. All of the extant conifers, such as Araucaria, cedar, celery-pine, cypress, fir, juniper, kauri, larch, pine, redwood, spruce, and yew, are included here. Some fossil conifers, however, belong to other distinct orders within the division Pinophyta. Multiple molecular studies indicate this order being paraphyletic with respect to Gnetales, with studies recovering Gnetales as either a sister group to Pinaceae or being more derived than Pinaceae but sister to the rest of the group. Brown (1825) first discerned that there were two groups of seed plants, distinguished by the form of seed development, based on whether the ovules were exposed, receiving pollen directly, or enclosed, which do not. Shortly afterwards, Brongniart (1828) coined the term Phanérogames gymnosperms to describe the former group. The distinction was then formalized by Lindley (1830), dividing what he referred to as the subclass Dicotyledons into two tribes, Gymnosperms and Angiosperms. In the gymnosperms (or Gymnospermae) Lindley included two orders, the Cycadeae and the Coniferae. In his final work (1853) he described Gymnogens as a class with four orders; Cycadeaceae (cycads) Pinaceae (conifers) Taxaceae (taxads) Gnetaceae In contrast, Bentham and Hooker (1880) included only three orders in the class Gymnospermeae, by including taxads within Coniferae; Gnetaceae Coniferae Cycadaceae In the Engler system (1903) Gymnospermae is listed as a subdivision (Unterabteilung) and adopted more of a splitter approach, including extinct taxa, with the following six classes; Cycadales Bennettitales Cordaitales Ginkgoales Coniferae Gnetales During this period, Gorozhankin published his treatise on Gymnosperms (1895), for which he bears the botanical authority for Pinales, Gorozh..

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Gnetophyta

Gnetophyta (nɛˈtɒfᵻtə,_ˈnɛtoʊfaɪtə) is a division of plants (alternatively considered the subclass Gnetidae or order Gnetales), grouped within the gymnosperms (which also includes conifers, cycads, and ginkgos), that consists of some 70 species across the three relict genera: Gnetum (family Gnetaceae), Welwitschia (family Welwitschiaceae), and Ephedra (family Ephedraceae). Fossilized pollen attributed to a close relative of Ephedra has been dated as far back as the Early Cretaceous.

Gymnosperm

The gymnosperms (dZIm'noUsp@rmz lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, Ginkgo, and gnetophytes, forming the clade Gymnospermae. The term gymnosperm comes from the composite word in γυμνόσπερμος (γυμνός and σπέρμα), literally meaning 'naked seeds'. The name is based on the unenclosed condition of their seeds (called ovules in their unfertilized state). The non-encased condition of their seeds contrasts with the seeds and ovules of flowering plants (angiosperms), which are enclosed within an ovary.

Pinaceae

The Pinaceae (pI'neIsi:%i:,_-si%aI), or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, piñons, larches, pines and spruces. The family is included in the order Pinales, formerly known as Coniferales. Pinaceae are supported as monophyletic by their protein-type sieve cell plastids, pattern of proembryogeny, and lack of bioflavonoids.

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