Land vertebrate faunachrons (LVFs) are biochronological units used to correlate and date terrestrial sediments and fossils based on their tetrapod faunas. First formulated on a global scale by Spencer G. Lucas in 1998, LVFs are primarily used within the Triassic Period (252 - 201 Ma), though Lucas later designated LVFs for other periods as well. Eight worldwide LVFs are defined for the Triassic. The first two earliest Triassic LVFs, the Lootsbergian and Nonesian, are based on South African synapsids and faunal assemblage zones estimated to correspond to the Early Triassic. These are followed by the Perovkan and Berdyankian, based on temnospondyl amphibians and Russian assemblages estimated to be from the Middle Triassic. The youngest four Triassic LVFs, the Otischalkian, Adamanian, Revueltian, and Apachean, are based on aetosaur and phytosaur reptiles common in the Late Triassic of the southwestern United States.
The LVF system, though widely used, is also a controversial application of biostratigraphy, as many Triassic tetrapods are rife with complications which endanger their utility as index fossils. Limited occurrences, inaccurate age estimates, overlapping LVF faunas, or taxonomic disagreement may jeopardize global correlations between Triassic tetrapods. This could render some LVFs as misleading assessments of Triassic faunal change through time. Regardless, Late Triassic phytosaurs are considered to have strong biostratigraphic utility even among detractors of Lucas's system.
Tetrapod biostratigraphy has been used for the Triassic of South Africa since 1906 and Argentina since 1966, but without much connection to global faunas. Starting in 1993, New Mexico Museum of Natural History and Science paleontologist Spencer G. Lucas and his colleagues began to define tetrapod biostratigraphy intervals in the Triassic of China and eastern and western North America. These named biostratigraphic intervals were inspired by the Land Mammal Age (LMA) system already in use for Cenozoic faunal assemblages.
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Typothorax is an extinct genus of typothoracine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: T. coccinarum, the type species, and T. antiquum. Typothorax was an aetosaur, a pseudosuchian distantly related to modern crocodilians. Unlike modern crocodilians, aetosaurs were herbivorous. Typothorax and other aetosaurs possess small, leaf-shaped teeth that were unsuited for a diet consisting of meat. Unlike some aetosaurs such as Desmatosuchus, Typothorax does not have large shoulder spikes.
Synapsids are one of the two major clades of vertebrate animals that evolved from basal amniotes, the other being the sauropsids, which include reptiles (turtles, crocodilians and lepidosaurs) and birds. The synapsids were once the dominant land animals in the late Paleozoic and early Mesozoic, but the only extant group that survived into the Cenozoic are the mammals. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name.
Since their appearance in the Anisian (ca. 240 Ma) scleractinian corals rapidly diversified and expandedacross shallow marine environments, becoming the main builders of reefs. This ecological success coralsowe to their symbiotic relationship with photosyn ...
Background: Scleractinian corals are currently a focus of major interest because of their ecological importance and the uncertain fate of coral reefs in the face of increasing anthropogenic pressure. Despite this, remarkably little is known about the evolu ...