This lecture explores how the brain specifies movements of multiple body parts with differing ranges of motion and reference frames to construct goal-directed behaviors. The instructor presents a surprising network architecture underlying coordinated movements, investigating neural ensemble recordings, perturbations, and modeling in mouse innate behaviors. Topics include the hierarchical view of movement specification, the role of the superior colliculus in encoding head and eye movement vectors, and the brainstem modules generating vectorial head movements and goal-directed saccades.