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This lecture explores the impact of climate change and permafrost thaw on Arctic and Boreal Ecosystems. It covers observed climate change in high latitudes, projected temperature changes, permafrost degradation, and the complex feedback mechanisms involving vegetation, organic matter decay, and greenhouse gas emissions. The instructor discusses the net ecosystem carbon balance, pathways of permafrost degradation, and the challenges in constraining permafrost feedback. Additionally, the lecture delves into the paleo perspective of permafrost and vegetation changes, highlighting past climate events. The presentation concludes by outlining the implications of permafrost collapse, soil carbon release, and the need for further research in this critical field.