The Tashtyk culture was Late Iron Age archaeological culture that flourished in the Yenisei valley in Siberia from the 1st century CE to the 4th century CE. Located in the Minusinsk Depression, environs of modern Krasnoyarsk, eastern part of Kemerovo Oblast, it was preceded by the Tagar culture. The Tashtyk culture was first surveyed by the Russian archaeologist Sergei Teploukhov. Teploukhov suggested that it had been initially Indo-European dominated, only to become overcome by the Yenisei Kirghiz around the 3rd century AD. The Yenisei Kirghiz are often associated with the Tashtyk culture. Tashtyk settlements and hill-forts have been unearthed throughout the Yenisei region, particularly the Sayan canyon area. Their most imposing monuments were immense barrows-crypt structures; these have yielded large quantities of clay and metal vessels and ornaments. In addition, numerous petrographic carvings have been found. Some of the graves contained leather models of human bodies with their heads wrapped in tissue and brightly painted. Inside the models there were small leather bags probably symbolising the stomach and containing burned human bones. Scaled-down replicas of swords, arrows and quivers were placed nearby. The animal motifs of the Tashtyk belonged to the Scytho-Altaic style, while they were also under significant Chinese influence. During his excavations of the Oglahty cemetery south of Minusinsk, Leonid Kyzlasov discovered a number of mummies with richly decorated plaster funerary masks showing Western Eurasian features, though this would not rule out some East Asian admixture, as revealed by ancient DNA (see below). There were also intact fur hats, silk clothes, and footwear (now in the Hermitage Museum, St. Petersburg). Oglakhty is considered as "the key site of stage I of the Tashtyk culture." From the early 20th century, various dates have been proposed for the Tashtyk burials: 1st century BC-1st century CE, 1st-2nd century CE. From the 1990s, new proposals were made dating the Tashtyk burials to the 3rd–4th centuries CE.