Forest dynamics

Forest dynamics describes the underlying physical and biological forces that shape and change a forest ecosystem. The continuous state of change in forests can be summarized with two basic elements: disturbance and succession. Disturbance (ecology) Forest disturbances are events that cause change in the structure and composition of a forest ecosystem, beyond the growth and death of individual organisms. Disturbances can vary in frequency and intensity, and include natural disasters such as fire, landslides, wind, volcanic eruptions, rare meteor impacts, outbreaks of insects, fungi, and other pathogens, animal-caused effects such as grazing and trampling, and anthropogenic disturbances such as warfare, logging, pollution, the clearing of land for urbanization or agriculture, and the introduction of invasive species. Not all disturbances are destructive or negative to the overall forest ecosystem. Many natural disturbances allow for renewal and growth and often release necessary nutrients. Small-scale disturbances are the key to creating and maintaining diversity and heterogeneity within a forest. Small-scale disturbances are events such as single-tree blowdowns, which create gaps that let light through the canopy to the understory and forest floor. This available light allows early-successional shade-intolerant species to colonize and maintain a population within the dominant forest, leading to the complex spatial mosaic forest structure recognized as old-growth. This process is referred to as patch dynamics or gap dynamics and has been described across many types of forests, including tropical, temperate, and boreal. The sets and patterns of natural disturbances that characterize a particular area or ecosystem are referred to as the ecosystem's disturbance regime. A natural community is closely linked with its natural disturbance regime. For example, temperate and boreal rainforests typically have a disturbance regime consisting of high-frequency but small-scale events, resulting in a highly complex forest dominated by very old trees.