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A mammoth is any species of the extinct elephantid genus Mammuthus. The various species of mammoth were commonly equipped with long, curved tusks. They lived from the Pliocene epoch (from around 5 million years ago) into the Holocene at about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are more closely related to living Asian elephants than African elephants. The oldest mammoth representative, Mammuthus subplanifrons, appeared around five million years ago during the early Pliocene in what is now southern and eastern Africa. Later in the Pliocene, by about three million years ago, mammoths dispersed into Eurasia, eventually covering most of Eurasia before migrating into North America around 1.5–1.3 million years ago, becoming ancestral to the Columbian mammoth (M. columbi). The last species to emerge, the woolly mammoth (M. primigenius), developed about 400,000 years ago in East Asia, with some surviving on Russia's Wrangel Island in the Arctic Ocean, as well as possibly the Taymyr Peninsula on mainland Siberia, until as recently as roughly 3,700 to 4,000 years ago, still extant during the existence of the earliest civilisations in Ancient Egypt and Mesopotamia. The earliest known proboscideans, the clade that contains the elephants, existed about 55 million years ago around the Tethys Sea area. The closest relatives of the Proboscidea are the sirenians and the hyraxes. The family Elephantidae is known to have existed six million years ago in Africa, and includes the living elephants and the mammoths. Among many now extinct clades, the mastodon is only a distant relative of the mammoths, and part of the separate Mammutidae family, which diverged 25 million years before the mammoths evolved. The following cladogram shows the placement of the genus Mammuthus among other proboscideans, based on hyoid characteristics: Since many remains of each species of mammoth are known from several localities, it is possible to reconstruct the evolutionary history of the genus through morphological studies.

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Woolly mammoth

The woolly mammoth (Mammuthus primigenius) is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with the African Mammuthus subplanifrons in the early Pliocene. The woolly mammoth began to diverge from the steppe mammoth about 800,000 years ago in East Asia. Its closest extant relative is the Asian elephant. The Columbian mammoth (Mammuthus columbi) lived alongside the woolly mammoth in North America, and DNA studies show that the two hybridised with each other.

Pleistocene

The Pleistocene (ˈplaɪstəˌsiːn,_-stoʊ- ; often referred to colloquially as the Ice Age) is the geological epoch that lasted from 2.58 million to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period.

Quaternary extinction event

The latter half of the Late Pleistocene to the beginning of the Holocene (~50,000-10,000 years Before Present) saw extinctions of numerous predominantly megafaunal species, which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by the widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence.

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