Hyrcania (hərˈkeɪniə) (Ὑρκανία Hyrkania, Old Persian: 𐎺𐎼𐎣𐎠𐎴 Varkâna, Middle Persian: 𐭢𐭥𐭫𐭢𐭠𐭭 Gurgān, Akkadian: Urqananu) is a historical region composed of the land south-east of the Caspian Sea in modern-day Iran and Turkmenistan, bound in the south by the Alborz mountain range and the Kopet Dag in the east. The region served as a satrapy (province) of the Median Empire, a sub-province of the Achaemenid Empire, and a province within its successors, the Seleucid, Arsacid and Sasanian empires. Hyrcania bordered Parthia to the east (later known as Abarshahr), Dihistan to the north, Media to the south and Mardia to the west. Hyrcania (Ὑρκανία) is the Greek name for the region, a borrowing from the Old Persian Verkâna as recorded in Darius the Great's Behistun Inscription (522 BC), as well as in other Old Persian cuneiform inscriptions. Verkā means "wolf" in Old Iranian, cf. Avestan vəhrkō, Gilaki and Mazandarani verk/verg, Modern Persian gorg, and Sanskrit vŗka (वृक). Consequently, Hyrcania means "Wolf-land". The name was extended to the Caspian Sea and underlies the name of the city Sari (Zadracarta), the then-largest city and the capital of ancient Hyrcania. Another archaic name, Dahistān (not to be confused with dehestan – a modern Iranian word for "district" or "county") is sometimes used interchangeably with Hyrcania. Dahistān refers, strictly speaking to the "place of the Dahae": an extinct people who lived immediately north of Hyrcania, as early as the 5th century BC. Hyrcania formed part of the Median Empire by 600 BC, and according to Nicolaus Damascenus, was administered as a satrapy by the time of the last Median king, Astyages, who appointed a certain Artasyras as satrap (governor). Upon the fall of the Median Empire, the region willingly submitted to the Achaemenid Empire and was occupied by Cyrus the Great in 549-548 BC, and for a time Artasyras continued as satrap under Cyrus, prior to his replacement by Astyages himself. According to Ctesias, Astyages' grandson Megabernes also served as satrap of Hyrcania.