A pyrena or pyrene (commonly called a "pit" or "stone") is the fruitstone within a drupe or drupelet produced by the ossification of the endocarp or lining of the fruit. It consists of a hard endocarp tissue surrounding one or more seeds (also called the "kernel"). The hardened endocarp which constitutes the pyrene provides a protective physical barrier around the seed, shielding it from pathogens and herbivory.
While many drupes are monopyrenous, containing only one pyrene, pome-type fruit with a hard, stony (rather than leathery) endocarp are typically polypyrenous drupes, containing multiple pyrenes.
The hardening of the endocarp of a developing drupe occurs via secondary cell wall formation and lignification. The biopolymer lignin, also found in wood, provides a structure within secondary cell walls which supports the polymerisation of cellulose and hemicellulose; together these polymers provide the endocarp with tensile strength and stiffness. Further hardening occurs during the biomineralisation of the endocarp. The biomineralisation of pyrenes during the life of the plant can aid the preservation of fruit remains in archaeological findings.
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Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. In some fruits, the edible portion is not derived from the ovary, but rather from the aril, such as the mangosteen or pomegranate, and the pineapple from which tissues of the flower and stem provide food. The grains of grasses are single-seed simple fruits wherein the pericarp and seed coat are fused into one layer.
The almond (Prunus amygdalus, syn. Prunus dulcis) is a species of small tree from the genus Prunus, cultivated worldwide for its seed, a culinary nut. Along with the peach, it is classified in the subgenus Amygdalus, distinguished from the other subgenera by corrugations on the shell (endocarp) surrounding the seed. The fruit of the almond is a drupe, consisting of an outer hull and a hard shell with the seed, which is not a true nut. Shelling almonds refers to removing the shell to reveal the seed.