The gentoo penguin (ˈdʒɛntuː ) (Pygoscelis papua) is a penguin species (or possibly a species complex) in the genus Pygoscelis, most closely related to the Adélie penguin (P. adeliae) and the chinstrap penguin (P. antarcticus). The earliest scientific description was made in 1781 by Johann Reinhold Forster with a type locality in the Falkland Islands. The species calls in a variety of ways, but the most frequently heard is a loud trumpeting, which the bird emits with its head thrown back. The application of "gentoo" to the penguin is unclear. Gentoo was an Anglo-Indian term to distinguish Hindus from Muslims. The English term may have originated from the Portuguese gentio ("pagan, gentile"). Some speculate that the white patch on the bird's head was thought to resemble a turban. It may also be a variation of another name for this bird, "Johnny penguin", with Johnny being Juanito in Spanish and sounds vaguely like gentoo. The Johnny rook, a predator, is likely named after the Johnny penguin. The specific name papua is a misnomer; in the original description, Johann Reinhold Forster, a naturalist who had circumnavigated the world with Captain James Cook, mistakenly assumed that the species occurred in Papua (New Guinea), the closest gentoos being over 6000 km to the south (on Macquarie Island). No penguins are found in New Guinea. Others trace the error to a "possibly fraudulent claim" in 1776 by French naturalist Pierre Sonnerat, who also alleged a Papuan location for the king penguin despite never having been to the island himself. The gentoo penguin is one of three species in the genus Pygoscelis. Mitochondrial and nuclear DNA evidence suggests the genus split from other penguins around 38 million years ago (Mya), about 2 million years after the ancestors of the genus Aptenodytes. In turn, the Adélie penguins split off from the other members of the genus around 19 Mya and the chinstrap and gentoo finally diverged around 14 Mya. There are 4 subspecies recognized by the International Ornithological Congress: P. p.