Concept

European shag

Summary
The European shag or common shag (Gulosus aristotelis) is a species of cormorant. It is the only member of the monotypic genus Gulosus. It breeds around the rocky coasts of western and southern Europe, southwest Asia and north Africa, mainly wintering in its breeding range except for the northernmost birds. In Britain this seabird is usually referred to as simply the shag. The scientific genus name derives from the Latin for glutton. The species name aristotelis commemorates the Greek philosopher Aristotle. The European shag was formerly classified within the genus Phalacrocorax, but a 2014 study found it to be significantly more diverged than the clade containing Phalacrocorax and Urile, but basal to the clade containing Nannopterum and Leucocarbo, and thus classified it in its own genus, Gulosus. The IOC followed this classification in 2021. Gulosus is thought to have split from the Nannopterum-Leucocarbo clade between 9.0 - 11.2 million years ago. There are three subspecies: G. a. aristotelis – (Linnaeus, 1761): nominate, found in northwestern Europe (Atlantic Ocean coasts) G. a. desmarestii – (Payraudeau, 1826): found in southern Europe, southwest Asia (Mediterranean and Black Sea coasts) G. a. riggenbachi – Hartert, 1923: found in northwest African coast The subspecies differ slightly in bill size and the breast and leg colour of young birds. Recent evidence suggests that birds on the Atlantic coast of southwest Europe are distinct from all three, and may be an as-yet undescribed subspecies. The name shag is also used in the Southern Hemisphere for several additional species of cormorants. This is a medium-large black bird, long and with a wingspan. It has a longish tail and yellow throat-patch. Adults have a small crest in the breeding season. It is distinguished from the great cormorant by its smaller size, lighter build, thinner bill, and, in breeding adults, by the crest and metallic green-tinged sheen on the feathers. Among those differences are that a shag has a lighter, narrower beak; and the juvenile shag has darker underparts.
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