Concept

Malawania

Malawania is an extinct genus of basal thunnosaur ichthyosaur known from the middle Early Cretaceous (Hauterivian or Barremian stage) of Iraq. Malawania was named by Valentin Fischer, Robert M. Appleby, Darren Naish, Jeff Liston, Riding, J. B., Brindley, S. and Pascal Godefroit in 2013 and the type species is Malawania anachronus. It is unusual as it is much more primitive than other Cretaceous ichthyosaurs, being most closely related to Ichthyosaurus from the Late Triassic and Early Jurassic, over 70 million years earlier than Malawania, with all other known ichthyosaurs from the Late Jurassic onwards belonging to the family Ophthalmosauridae. The holotype and only known specimen was discovered in 1952 by British petroleum geologists D. M. Morton, F. R. S. Henson, R. J. Wetzel and L. C. F. Damesin while working in Chia Gara, Amadia in Iraqi Kurdistan. The slab was being used to dam a small river and was part of a mule track. It was then transported back to the UK and donated to the Natural History Museum in 1959. Robert M. Appleby would study the specimen for many years until his death in 2004, but never published a paper, one manuscript submitted to Paleontology in 1979 was rejected due to the uncertain provenance of the specimen. The specimen was formally described in 2013 by Fischer and colleagues. Using palynology it was determined that the rocks surrounding the specimen were Hauterivian-Barremian in age, which was unexpected given the archaic nature of the specimen. Each lacrimal (paired bones at the front edges of the orbits) bears a hollow towards its front edge. Fischer and colleagues speculated that these may have been the location of the lacrimal glands (which produce the tear film). The long, backwards projecting processes of the lacrimals extend halfway underneath the orbits (eye sockets). The eyeballs were supported by rings of 13 bones, known as sclerotic rings. Malawania has a minimum of five neck vertebrae. While the boundary between them is still demarcated, the first two vertebrae are fused.

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