Lobopodia

Lobopodians are members of the informal group Lobopodia (from the Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998). They are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, marine worm-like fossil panarthropods such as Aysheaia and Hallucigenia. The oldest near-complete fossil lobopodians date to the Lower Cambrian; some are also known from Ordovician, Silurian and Carboniferous Lagerstätten. Some bear toughened claws, plates or spines, which are commonly preserved as carbonaceous or mineralized microfossils in Cambrian strata. The grouping is considered to be paraphyletic, as the three living panarthropod groups (Arthropoda, Tardigrada and Onychophora) are thought to have evolved from lobopodian ancestors. The Lobopodian concept varies from author to author. Its most general sense refers to a suite of mainly Cambrian worm-like panarthropod taxa possessing lobopods – for example, Aysheaia, Hallucigenia, and Xenusion – which were traditionally united as "Xenusians" or "Xenusiids" (class Xenusia). Certain Dinocaridid genera, such as Opabinia, Pambdelurion, and Kerygmachela, may also be regarded as lobopodians, sometimes referred to more specifically as "gilled lobopodians" or "gilled lobopods". This traditional, informal usage of "Lobopodia" treats it as an evolutionary grade, including only extinct Panarthropods near the base of crown Panarthropoda. Crown Panarthropoda comprises the three extant Panarthropod phyla – Onychophora (velvet worms), Tardigrada (waterbears), and Arthropoda (arthropods) – as well as their most recent common ancestor and all of its descendants. Thus, in this usage, Lobopodia consists of various basal Panarthropods. This corresponds to "A" in the image to the left. An alternative, broader definition of Lobopodia would also incorporate Onychophora and Tardigrada, the two living panarthropod phyla which still bear lobopodous limbs.