Guard cells are specialized plant cells in the epidermis of leaves, stems and other organs that are used to control gas exchange. They are produced in pairs with a gap between them that forms a stomatal pore. The stomatal pores are largest when water is freely available and the guard cells become turgid, and closed when water availability is critically low and the guard cells become flaccid. Photosynthesis depends on the diffusion of carbon dioxide (CO2) from the air through the stomata into the mesophyll tissues. Oxygen (O2), produced as a byproduct of photosynthesis, exits the plant via the stomata. When the stomata are open, water is lost by evaporation and must be replaced via the transpiration stream, with water taken up by the roots. Plants must balance the amount of CO2 absorbed from the air with the water loss through the stomatal pores, and this is achieved by both active and passive control of guard cell turgor pressure and stomatal pore size. Guard cells are cells surrounding each stoma. They help to regulate the rate of transpiration by opening and closing the stomata. Light is the main trigger for the opening or closing. Each guard cell has a relatively thick cuticle on the pore-side and a thin one opposite it. As water enters the cell, the thin side bulges outward like a balloon and draws the thick side along with it, forming a crescent; the combined crescents form the opening of the pore. Guard cells contain phototropin proteins which are serine and threonine kinases with blue-light photoreceptor activity. Phototrophins contain two light, oxygen, and voltage sensor (LOV) domains, and are part of the PAS domain superfamily. The phototropins trigger many responses such as phototropism, chloroplast movement and leaf expansion as well as stomatal opening. Not much was known about how these photoreceptors worked prior to around 1998. The mechanism by which phototropins work was elucidated through experiments with broad bean (Vicia faba).

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Related concepts (5)
Leaf
A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, flower, and fruit collectively form the shoot system. In most leaves, the primary photosynthetic tissue is the palisade mesophyll and is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral.
Transpiration
Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. Water is necessary for plants but only a small amount of water taken up by the roots is used for growth and metabolism. The remaining 97–99.5% is lost by transpiration and guttation. Leaf surfaces are dotted with pores called stomata (singular "stoma"), and in most plants they are more numerous on the undersides of the foliage.
Turgor pressure
Turgor pressure is the force within the cell that pushes the plasma membrane against the cell wall. It is also called hydrostatic pressure, and is defined as the pressure in a fluid measured at a certain point within itself when at equilibrium. Generally, turgor pressure is caused by the osmotic flow of water and occurs in plants, fungi, and bacteria. The phenomenon is also observed in protists that have cell walls. This system is not seen in animal cells, as the absence of a cell wall would cause the cell to lyse when under too much pressure.
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