Odonata is an order of flying insects that includes the dragonflies and damselflies. Members of the group first appeared during the Triassic, though members of their total group, Odonatoptera, first appeared in Late Carboniferous. The two common groups are distinguished with dragonflies, placed in the suborder Epiprocta, usually being larger, with eyes together and wings up or out at rest, while damselflies, suborder Zygoptera, are usually smaller with eyes placed apart and wings along body at rest. All Odonata have aquatic larvae called naiads (nymphs), and all of them, larvae and adults, are carnivorous. The adults can land, but rarely walk. Their legs are specialised for catching prey. They are almost entirely insectivorous. Fabricius coined the term Odonata in 1793 from the Ancient Greek ὀδών (Ionic form of ὀδούς ) 'tooth'. One hypothesis is that it was because their maxillae are notably toothed. Most insects also have toothed mandibles. The word dragonfly usually refers to only Anisoptera, but is also sometimes used to refer to all Odonata Odonata enthusiasts avoid ambiguity by using the term true dragonfly, or simply anisopteran, when referring to just the Anisoptera. The term warriorfly has also been proposed. Some 7,000 species have been described in this order. This order is closely related to mayflies and several extinct orders in a group called the Palaeoptera, but this grouping might be paraphyletic. What they do share with mayflies is the nature of how the wings are articulated and held in rest (see insect flight for a detailed discussion). In some treatments, the Odonata are understood in an expanded sense, essentially synonymous with the superorder Odonatoptera but not including the prehistoric Protodonata. In this approach, instead of Odonatoptera, the term Odonatoidea is used. The systematics of the "Palaeoptera" are by no means resolved; what can be said however is that regardless of whether they are called "Odonatoidea" or "Odonatoptera", the Odonata and their extinct relatives do form a clade.
Pedro Miguel Nunes Pereira de Almeida Reis, Matteo Pezzulla, Yuexia Luna Lin