Summary
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (front to back), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm. The most commonly cited functions of the notochord are: as a midline staple that provides directional reference to surrounding tissue during embryonic development, as an axial endoskeleton (structural element) and vertebral precursor, and as an elastic spring that allows more efficient tail motion when swimming. In lancelets, the notochord persists throughout life as the main structural support of the body. In tunicates the notochord is present only in the larval stage, being completely absent in the adult animal. In these invertebrate chordates, the notochord is not vacuolated. In all vertebrates other than the hagfish, the notochord is integrated into the vertebral column, with its original structure being (nearly) retained in the intervertebral discs as the nucleus pulposus. The notochord is a long, rod-like midline structure that develops dorsal to the gut tube and ventral to the neural tube. The notochord is composed primarily of a core of glycoproteins encased in a sheath of collagen fibers wound into two opposing helices. The glycoproteins are stored in vacuolated, turgid cells, which are covered with caveolae on their cell surface. The angle between these fibers determines whether increased pressure in the core will result in shortening and thickening versus lengthening and thinning. Alternating contraction of muscle fibers attached to each side of the notochord result in a side-to-side motion resembling stern sculling, which allows tail swimming and undulation. The stiffened notochord prevents movement through telescoping motion such as that of an earthworm.
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Related concepts (61)
Haikouichthys
Haikouichthys ˌhaɪkuˈɪkθɪs is an extinct genus of craniate (animals with notochords and distinct heads) that lived 518 million years ago, during the Cambrian explosion of multicellular life. Haikouichthys had a defined skull and other characteristics that have led paleontologists to label it a true craniate, and even to be popularly characterized as one of the earliest fishes. Cladistic analysis indicates that the animal is probably a basal chordate or a basal craniate; but it does not possess sufficient features to be included uncontroversially even in either stem group.
Notochord
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (front to back), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm.
Vertebral column
The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordates has been replaced by a segmented series of bone: vertebrae separated by intervertebral discs. Individual vertebrae are named according to their region and position, and can be used as anatomical landmarks in order to guide procedures such as lumbar punctures.
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