The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina.
It comprises nine living species in two genera, the type of which is the genus Nautilus. Though it more specifically refers to species Nautilus pompilius, the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between .
Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle. Having survived relatively unchanged for hundreds of millions of years, nautiluses represent the only living members of the subclass nautiloidea, and are often considered "living fossils".
The word nautilus is derived from the Greek word ναυτίλος nautílos "sailor", it originally referred to a type of octopus of the genus Argonauta, also known as 'paper nautilus', which were thought to use two of their arms as sails.
The "tentacles" of the nautilus are actually cirri (singular: cirrus), composed of long, soft, flexible appendages which are retractable into corresponding hardened sheaths. Unlike the 8–10 head appendages of coleoid cephalopods, nautiluses have many cirri. In the early embryonic stages of nautilus development, a single molluscan foot differentiates into a total of 60–90 cirri, varying even within a species. Nautilus cirri also differ from the tentacles of some coleoids in that they are non-elastic and lack pads or suckers. Instead, nautilus cirri adhere to prey using their ridged surface. Nautiluses have a powerful grip, and attempts to take an object already grasped by a nautilus may tear away the animal's cirri, which will remain firmly attached to the surface of the object.
The main cirri emerge from sheaths which cohere into a single firm fleshy mass.