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This lecture covers the fundamental concepts of file handling in C, including opening, reading, writing, and closing files. It also delves into the intricacies of endianness, explaining how data is ordered in memory and on disk. The lecture emphasizes the importance of understanding the representation of numbers and the challenges posed by different conventions. Additionally, it explores the significance of characters like the end-of-line character and provides insights into file paths, current paths, and file access errors. The session concludes with a quiz on file handling and endianness.