This lecture discusses the mechanisms of photosynthesis, focusing on both oxygenic and anoxygenic processes. It begins with an overview of the importance of photosynthesis in microbial ecosystems, particularly in wastewater treatment and carbon capture technologies. The instructor explains the historical context of photosynthesis, detailing the evolution of prokaryotic organisms and their roles in altering Earth's atmosphere. Key concepts include the differences between oxygenic and anoxygenic photosynthesis, the types of pigments involved, and the structures that facilitate light capture. The lecture also covers the ecological arrangements of photosynthetic organisms in various environments, highlighting the stratification of microbial mats. The instructor emphasizes the significance of understanding these processes in addressing contemporary environmental challenges, such as CO2 emissions and the development of sustainable technologies. The session concludes with a discussion on the evolutionary origins of photosynthesis and its implications for higher life forms.