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This lecture covers the structure, function, and regulation of microtubules, focusing on topics such as the three principal cytoskeletal polymers, microtubule organization by centrioles, microtubule assembly, accessory proteins, motor proteins, and the forces involved in microtubule dynamics. The instructor explains the role of small compounds in modulating microtubules, force generation through polymerization, and the mechanochemical cycle of motor proteins. Additionally, the lecture delves into the process of pronuclear migration, highlighting the forces involved and the mechanisms of genome unification.