Trigonotarbida

The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian (Pridoli to Sakmarian). These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton (tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tubercles. About seventy species are currently known, with most fossils originating from the Carboniferous coal measures. The first trigonotarbid was described in 1837 from the coal measures of Coalbrookdale in England by the famous English geologist Dean William Buckland. He believed it to be a fossil beetle and named it Curculoides prestvicii. A much better preserved example was later discovered from Coseley near Dudley; also in the English West Midlands conurbation. Described in 1871 by Henry Woodward, he correctly identified it as an arachnid and renamed it Eophrynus prestvicii—whereby the genus name comes from ἠώς (eos, meaning 'dawn'), and Phrynus, a genus of living whip spider (Amblypygi). Woodward subsequently described another trigonotarbid, Brachypyge carbonis, from the coal measures of Mons in Belgium; although this fossil is known only from its abdomen and was initially mistaken for those of a crab. In 1882, the German zoologist Ferdinand Karsch described a number of fossil arachnids from the coal measures of Neurode in Silesia (now Poland), including one he named Anthracomartus voelkelianus in honour of Herr Völkel, the foreman of the mine where it was discovered. This species was raised to a new, extinct, arachnid order which Karsch called Anthracomarti.