Concept

Sap

Summary
Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separate substance, separately produced, and with different components and functions. Insect honeydew is called sap, particularly when it falls from trees, but is only the remains of eaten sap and other plant parts. Saps may be broadly divided into two types: xylem sap and phloem sap. Xylem sap (pronounced ˈzaɪləm) consists primarily of a watery solution of hormones, mineral elements and other nutrients. Transport of sap in xylem is characterized by movement from the roots toward the leaves. Over the past century, there has been some controversy regarding the mechanism of xylem sap transport; today, most plant scientists agree that the cohesion-tension theory best explains this process, but multiforce theories that hypothesize several alternative mechanisms have been suggested, including longitudinal cellular and xylem osmotic pressure gradients, axial potential gradients in the vessels, and gel- and gas-bubble-supported interfacial gradients. Xylem sap transport can be disrupted by cavitation—an "abrupt phase change [of water] from liquid to vapor"—resulting in air-filled xylem conduits. In addition to being a fundamental physical limit on tree height, two environmental stresses can disrupt xylem transport by cavitation: increasingly negative xylem pressures associated with water stress, and freeze-thaw cycles in temperate climates. Phloem sap (pronounced ˈfloʊɛm) consists primarily of sugars, hormones, and mineral elements dissolved in water. It flows from where carbohydrates are produced or stored (sugar source) to where they are used (sugar sinks). The pressure flow hypothesis proposes a mechanism for phloem sap transport, although other hypotheses have been proposed. Phloem sap is thought to play a role in sending informational signals throughout vascular plants.
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