Concept

Wiwaxia

Summary
Wiwaxia is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. Wiwaxia fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to 5 cm (2 inch) when fully grown, although a range of juvenile specimens are known, the smallest being long. Wiwaxia's affinity has been a matter of debate: Researchers were long split between two possibilities. On the one hand, its rows of scales looked superficially similar to certain scale worms (annelids); conversely, its mouthparts and general morphology suggested a relationship to the shell-less molluscs. More recently, evidence for a molluscan affinity has been accumulating, based on new details of Wiwaxias mouthparts, scales, and growth history. The proposed clade Halwaxiida contains Wiwaxia as well as several similar Cambrian animals. This article concentrates on the species Wiwaxia corrugata, which is known from hundreds of complete specimens in the Burgess Shale; other species are known only from fragmentary material or limited sample sizes. Wiwaxia was bilaterally symmetrical; viewed from the top the body was elliptical with no distinct head or tail, and from the front or rear it was almost rectangular. It reached in length. Estimating their height is difficult because specimens were compressed after death; a typical specimen may have been high excluding the spines on their backs. The ratio of width to length does not appear to change as the animals grew. Wiwaxias flat underside was soft and unarmored; most of the surface was occupied by a single slug-like foot. Little is known of the internal anatomy, although the gut apparently ran straight and all the way from the front to the rear. At the front end of the gut, about from the animal's front in an average specimen about long, there was a feeding apparatus that consisted of two (or in rare large specimens three) rows of backward-pointing conical teeth.
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