CrocodiliaCrocodilia (or Crocodylia, both krQk@'dIli@) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 94 million years ago in the Late Cretaceous period (Cenomanian stage) and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the order's total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period, and diversified during the Mesozoic era.
TomistomaTomistoma is a genus of gavialid crocodilians. They are noted for their long narrow snouts used to catch fish, similar to the gharial. Tomistoma contains one extant (living) member, the false gharial (Tomistoma schlegelii), as well as potentially several extinct species: T. cairense, T. lusitanicum T. coppensi, and T. dowsoni. However, these species may need to be reclassified to different genera as studies have shown them to be paraphyletic, for example: previously assigned species T.
False gharialThe false gharial (Tomistoma schlegelii), also known by the names Malayan gharial, Sunda gharial and tomistoma is a freshwater crocodilian of the family Gavialidae native to Peninsular Malaysia, Borneo, Sumatra and Java. It is listed as Vulnerable on the IUCN Red List, as the global population is estimated at around 2,500 to 10,000 mature individuals. The specific name schlegelii honors Hermann Schlegel. The scientific name Crocodilus (Gavialis) schlegelii was proposed by Salomon Müller in 1838 who described a specimen collected in Borneo.
GavialisGavialis is a genus of crocodylians that includes the living gharial Gavialis gangeticus and one known extinct species, Gavialis bengawanicus. G. gangeticus comes from the Indian Subcontinent, while G. bengawanicus is known from Java. Gavialis likely first appeared in the Indian Subcontinent in the Pliocene and dispersed into the Malay Archipelago through a path called the Siva–Malayan route in the Quaternary.
IkanogavialisIkanogavialis is an extinct genus of gavialid crocodilian. Fossils have been found in the Urumaco Formation in Urumaco, Venezuela and the Solimões Formation of Brazil. The strata from which remains are found are late Miocene in age, rather than Pliocene as was once thought. A possible member of this genus survived into the Late Holocene on Muyua or Woodlark Island in Papua New Guinea. Ikanogavialis had a dorsoventrally deep snout and a distinctive notch between the dentary and maxillary alveoli.
GavialosuchusGavialosuchus is an extinct genus of gavialoid crocodylian from the early Miocene of Europe. Currently only one species is recognized, as a few other species of Gavialosuchus have since been reclassified to other genera. The type species, G. eggenburgensis, is known from the early Miocene of Austria. Two other species - G. americanus and G. carolinensis - have since been reclassified to other genera. Myrick Jr. (2001) proposed synonymizing Gavialosuchus americanus with Thecachampsa antiqua. Piras et al.
HarpacochampsaHarpacochampsa is a poorly known Early Miocene crocodilian from the Bullock Creek lagerstätte of the Northern Territory, Australia. The current specimen consists of a partial skull and fragments of a long, slender snout reminiscent of that of a false gharial, demonstrating that it was a piscivore in life. It was originally tentatively placed within a group of Australian crocodilians now known as the Mekosuchinae, although this has been frequently disputed, with other authors instead suggesting it may have been a more basal crocodyloid or a type of gavialid.
GryposuchusGryposuchus is an extinct genus of gavialid crocodilian. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the Miocene epoch (Colhuehuapian to Huayquerian). One recently described species, G. croizati, grew to an estimated length of . Gryposuchus is the type genus of the subfamily Gryposuchinae, although a 2018 study indicates that Gryposuchinae and Gryposuchus might be paraphyletic and rather an evolutionary grade towards the gharial.
PiscogavialisPiscogavialis is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is P. jugaliperforatus. Fossils of Piscogavialis have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998, where it coexisted with the much smaller gavialid Sacacosuchus. Piscogavialis is known only from a single specimen, but it represents some of the best preserved gavialid material known from South America. The skull is preserved in three dimensions and is nearly complete.
PenghusuchusPenghusuchus is an extinct genus of gavialid crocodylian. It is known from a skeleton found in Middle to Upper Miocene rocks of Penghu Island, off Taiwan. The taxon was described in 2009 by Shan and colleagues; the type species is P. pani. It may be related to two other fossil Asian gavialids: Toyotamaphimeia machikanensis of Japan and Hanyusuchus sinensis of South China. It was a medium-sized gavialid with an estimated total length of . On 25 March 2006, on the coast of Neian, Shiyu, Penghu Islands, an excavator driver Mr.
