Concept

False gharial

Summary
The false gharial (Tomistoma schlegelii), also known by the names Malayan gharial, Sunda gharial and tomistoma is a freshwater crocodilian of the family Gavialidae native to Peninsular Malaysia, Borneo, Sumatra and Java. It is listed as Vulnerable on the IUCN Red List, as the global population is estimated at around 2,500 to 10,000 mature individuals. The specific name schlegelii honors Hermann Schlegel. The scientific name Crocodilus (Gavialis) schlegelii was proposed by Salomon Müller in 1838 who described a specimen collected in Borneo. In 1846, he proposed to use the name Tomistoma schlegelii, if it needs to be placed in a distinct genus. The genus Tomistoma potentially also contains several extinct species like T. cairense, T. lusitanicum, T. taiwanicus, and T. coppensi. However, these species may need to be reclassified to different genera as evidence suggests they may be paraphyletic. The false gharial's snout broadens considerably towards the base and so is more similar to those of true crocodiles than to the gharial (Gavialis gangeticus), whose osteology indicated a distinct lineage from all other living crocodilians. However, although more morphologically similar to Crocodylidae based on skeletal features, recent molecular studies using DNA sequencing consistently indicate that the false gharial and by inference other related extinct forms traditionally viewed as belonging to the crocodylian subfamily Tomistominae actually belong to Gavialoidea and Gavialidae. Fossils of extinct Tomistoma species have been found in deposits of Paleogene, Neogene, and Quaternary ages in Taiwan, Uganda, Italy, Portugal, Egypt and India, but nearly all of them are likely to be distinct genera due to older age compared to the false gharial. The below cladogram of the major living crocodile groups is based on molecular studies and shows the false gharial's close relationships: The following cladogram shows the false gharial's placement within the Gavialidae; it is based on a tip dating study, for which morphological, DNA sequencing and stratigraphic data were analysed: The false gharial is dark reddish-brown above with dark brown or black spots and cross-bands on the back and tail.
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