A keel or carina (plural carinae) in bird anatomy is an extension of the sternum (breastbone) which runs axially along the midline of the sternum and extends outward, perpendicular to the plane of the ribs. The keel provides an anchor to which a bird's wing muscles attach, thereby providing adequate leverage for flight. Not all birds have keels; in particular, some flightless birds lack a keel structure. Some flightless birds have a keel, such as the penguin; but in the penguin's case, its wings are too small for its body, so flight would require flapping its wings too fast to be practical.
Historically, the presence or absence of a pronounced keel structure was used as a broad classification of birds into two orders: Carinatae (from carina, "keel"), having a pronounced keel; and ratites (from ratis, "raft" - referring to the flatness of the sternum), having a subtle keel structure or lacking one entirely. However, this classification has become disused as evolutionary studies have shown that many flightless birds have evolved from flighted birds.
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Bird flight is the primary mode of locomotion used by most bird species in which birds take off and fly. Flight assists birds with feeding, breeding, avoiding predators, and migrating. Bird flight is one of the most complex forms of locomotion in the animal kingdom. Each facet of this type of motion, including hovering, taking off, and landing, involves many complex movements. As different bird species adapted over millions of years through evolution for specific environments, prey, predators, and other needs, they developed specializations in their wings, and acquired different forms of flight.
Flightless birds are birds that, through evolution, lost the ability to fly. There are over 60 extant species, including the well-known ratites (ostriches, emu, cassowaries, rheas, and kiwi) and penguins. The smallest flightless bird is the Inaccessible Island rail (length 12.5 cm, weight 34.7 g). The largest (both heaviest and tallest) flightless bird, which is also the largest living bird in general, is the ostrich (2.7 m, 156 kg).