Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner bark, which in older stems is living tissue, includes the innermost layer of the periderm. The outer bark on older stems includes the dead tissue on the surface of the stems, along with parts of the outermost periderm and all the tissues on the outer side of the periderm. The outer bark on trees which lies external to the living periderm is also called the rhytidome.
Products derived from bark include bark shingle siding and wall coverings, spices and other flavorings, tanbark for tannin, resin, latex, medicines, poisons, various hallucinogenic chemicals and cork. Bark has been used to make cloth, canoes, and ropes and used as a surface for paintings and map making. A number of plants are also grown for their attractive or interesting bark colorations and surface textures or their bark is used as landscape mulch.
The process of removing bark is decortication and a log or trunk from which bark has been removed is said to be decorticated.
Bark is present only on woody plants - herbaceous plants and stems of young plants lack bark.
From the outside to the inside of a mature woody stem, the layers include the following:
Bark
Periderm
Cork (phellem or suber), includes the rhytidome
Cork cambium (phellogen)
Phelloderm
Cortex
Phloem
Vascular cambium
Wood (xylem)
Sapwood (alburnum)
Heartwood (duramen)
Pith (medulla)
In young stems, which lack what is commonly called bark, the tissues are, from the outside to the inside:
Epidermis, which may be replaced by periderm
Cortex
Primary and secondary phloem
Vascular cambium
Secondary and primary xylem.
Cork cell walls contain suberin, a waxy substance which protects the stem against water loss, the invasion of insects into the stem, and prevents infections by bacteria and fungal spores.
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2023
In recent years, xylem sap composition has been shown to affect xylem hydraulics. However, information on how much xylem sap composition can vary across seasons and specifically under drought stress is still limited. We measured xylem sap chemical composit ...
Oxford2023
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