Swarm behaviour, or swarming, is a collective behaviour exhibited by entities, particularly animals, of similar size which aggregate together, perhaps milling about the same spot or perhaps moving en masse or migrating in some direction. It is a highly interdisciplinary topic. As a term, swarming is applied particularly to insects, but can also be applied to any other entity or animal that exhibits swarm behaviour. The term flocking or murmuration can refer specifically to swarm behaviour in birds, herding to refer to swarm behaviour in tetrapods, and shoaling or schooling to refer to swarm behaviour in fish. Phytoplankton also gather in huge swarms called blooms, although these organisms are algae and are not self-propelled the way animals are. By extension, the term "swarm" is applied also to inanimate entities which exhibit parallel behaviours, as in a robot swarm, an earthquake swarm, or a swarm of stars. From a more abstract point of view, swarm behaviour is the collective motion of a large number of self-propelled entities. From the perspective of the mathematical modeller, it is an emergent behaviour arising from simple rules that are followed by individuals and does not involve any central coordination. Swarm behaviour is also studied by active matter physicists as a phenomenon which is not in thermodynamic equilibrium, and as such requires the development of tools beyond those available from the statistical physics of systems in thermodynamic equilibrium. In this regard, swarming has been compared to the mathematics of superfluids, specifically in the context of starling flocks (murmuration). Swarm behaviour was first simulated on a computer in 1986 with the simulation program boids. This program simulates simple agents (boids) that are allowed to move according to a set of basic rules. The model was originally designed to mimic the flocking behaviour of birds, but it can be applied also to schooling fish and other swarming entities.

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Related concepts (21)
Shoaling and schooling
In biology, any group of fish that stay together for social reasons are shoaling, and if the group is swimming in the same direction in a coordinated manner, they are schooling. In common usage, the terms are sometimes used rather loosely. About one quarter of fish species shoal all their lives, and about one half shoal for part of their lives. Fish derive many benefits from shoaling behaviour including defence against predators (through better predator detection and by diluting the chance of individual capture), enhanced foraging success, and higher success in finding a mate.
Stigmergy
Stigmergy (ˈstɪɡmərdʒi ) is a mechanism of indirect coordination, through the environment, between agents or actions. The principle is that the trace left in the environment by an individual action stimulates the performance of a succeeding action by the same or different agent. Agents that respond to traces in the environment receive positive fitness benefits, reinforcing the likelihood of these behaviors becoming fixed within a population over time. Stigmergy is a form of self-organization.
Flocking
Flocking is the behavior exhibited when a group of birds, called a flock, are foraging or in flight. Computer simulations and mathematical models that have been developed to emulate the flocking behaviours of birds can also generally be applied to the "flocking" behaviour of other species. As a result, the term "flocking" is sometimes applied, in computer science, to species other than birds, to mean collective motion by a group of self-propelled entities, a collective animal behaviour exhibited by many living beings such as fish, bacteria, and insects.
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