The Columbian mammoth (Mammuthus columbi) is an extinct species of mammoth that inhabited the Americas as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from mammoths that colonised North America around 1.5 million years ago, that later hybridised with woolly mammoths during the Middle Pleistocene, prior to 420,000 years ago. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant. Reaching at the shoulders and in weight, the Columbian mammoth was one of the largest species of mammoth. It had long, curved tusks and four molars, which were replaced six times during the lifetime of an individual. It most likely used its tusks and trunk like modern elephants—for manipulating objects, fighting, and foraging. Bones, hair, dung, and stomach contents have been discovered, but no preserved carcasses are known. The Columbian mammoth preferred open areas, such as parkland landscapes, and fed on sedges, grasses, and other plants. It did not live in the Arctic regions of Canada, which were instead inhabited by woolly mammoths. The ranges of the two species may have overlapped, and genetic evidence suggests that they interbred. Several sites contain the skeletons of multiple Columbian mammoths, either because they died in incidents such as a drought, or because these locations were natural traps in which individuals accumulated over time. For a few thousand years prior to their extinction, Columbian mammoths coexisted in North America with Paleoindians – the first humans to inhabit the Americas – who hunted them for food, used their bones for making tools, and possibly depicted them in ancient art. Columbian mammoth remains have been found in association with Clovis culture artifacts; these remains stemmed from hunting as well as possibly scavenging.