Concept

Bombardier beetle

Summary
Bombardier beetles are ground beetles (Carabidae) in the tribes Brachinini, Paussini, Ozaenini, or Metriini—more than 500 species altogether—which are most notable for the defense mechanism that gives them their name: when disturbed, they eject a hot noxious chemical spray from the tip of the abdomen with a popping sound. The spray is produced from a reaction between two hypergolic chemical compounds, hydroquinone and hydrogen peroxide, which are stored in two reservoirs in the beetle's abdomen. When the aqueous solution of hydroquinones and hydrogen peroxide reaches the "vestibule" (Eisner's word), catalysts facilitate the decomposition of the hydrogen peroxide and the oxidation of the hydroquinone. Heat from the reaction brings the mixture to near the boiling point of water and produces gas that drives the ejection. The damage caused can be fatal to attacking insects. Some bombardier beetles can direct the spray in a wide range of directions. The beetle's unusual defense mechanism is claimed by some creationists to be an example of what they call irreducible complexity, though this is refuted by evolutionary biologists. Bombardier beetles inhabit all the continents except Antarctica. They typically live in woodlands or grasslands in the temperate zones but can be found in other environments if there are moist places to lay their eggs. Most species of bombardier beetles are carnivorous, including the larva. The beetle typically hunts at night for other insects, but will often congregate with others of its species when not actively looking for food. There are two large glands that open at the tip of the abdomen. Each gland is composed of a thick walled vestibule which contains a mixture of catalases and peroxidases produced by the secretory cells that line the vestibule. Both glands are also made up of a thin-walled and compressible reservoir which contains an aqueous solution of hydroquinones and hydrogen peroxide. When the beetle feels threatened it opens a valve which allows the aqueous solution from the reservoir to reach the vestibule.
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