Concept

Ornithoscelida

Ornithoscelida (ˌɔːrnɪθəˈsɛlɪdə) is a proposed clade that includes various major groupings of dinosaurs. An order Ornithoscelida was originally proposed by Thomas Henry Huxley but later abandoned in favor of Harry Govier Seeley's division of Dinosauria into Saurischia and Ornithischia. The term was revived in 2017 after a new cladistic analysis by Baron et al. Thomas Henry Huxley originally defined the term in an 1869 lecture as a group comprising two subgroups: the large and heavy-set Dinosauria and the newly discovered Compsognathus, which he placed in a new grouping Compsognatha. The former were defined by their shorter cervical vertebrae, and the femur length exceeding tibia length, and the latter with longer cervical vertebrae, and the femur length shorter than tibia length. He noted that the characteristics of their bones showed many features akin to birds. The dinosaurs Huxley had divided into three families: Megalosauridae: Teratosaurus, Palaeosaurus, Megalosaurus, Poekilopleuron, Laelaps, and Euskelosaurus (tentatively) Scelidosauridae: Thecodontosaurus, Scelidosaurus, Hylaeosaurus, Polacanthus (tentatively), and Acanthopholis Iguanodontidae: Cetiosaurus, Iguanodon, Hypsilophodon, Hadrosaurus, and Stenopelix (tentatively) S. Williston (1878) included Compsognathus in Dinosauria and divided dinosaurs into Sauropoda and Ornithoscelida, the latter including taxa that would later be considered theropods and ornithischians. This classification quickly fell out of use, due to the dominant classification system by Harry Govier Seeley that grouped dinosaurs into two main branches: Saurischia and Ornithischia. In the beginning of the twenty-first century, improved descriptions of the early ornithischians Heterodontosaurus and Lesothosaurus vastly increased the available information on the origins of the Ornithischia. In March 2017, a paper in the journal Nature by Matthew Baron, David Norman, and Paul Barrett, published an analysis in which the theropod dinosaurs — no longer containing the Herrerasauridae — were more closely related to ornithischian dinosaurs than to the Sauropodomorpha, the group to which the sauropod dinosaurs belong.

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