Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire.
Some lichens grow on rocks without soil, so may be among the first of life forms, and break down the rocks into soil for plants. Since some uninhabited land may have thin, poor quality soils with few nutrients, pioneer species are often hardy plants with adaptations such as long roots, root nodes containing nitrogen-fixing bacteria, and leaves that employ transpiration. Note that they are often photosynthetic plants, as no other source of energy (such as other species) except light energy is often available in the early stages of succession, thus making it less likely for a pioneer species to be non-photosynthetic.
The plants that are often pioneer species also tend to be wind-pollinated rather than insect-pollinated, as insects are unlikely to be present in the usually barren conditions in which pioneer species grow; however, pioneer species tend to reproduce asexually altogether, as the extreme or barren conditions present make it more favourable to reproduce asexually in order to increase reproductive success rather than invest energy into sexual reproduction. Pioneer species will eventually die, create plant litter, and break down as "leaf mold" after some time, making new soil for secondary succession (see below), and releasing nutrients for small fish and aquatic plants in adjacent bodies of water.
Some examples of pioneering plant species:
Barren sand - lyme grass (Leymus arenarius), sea couch grass (Agropyron pungens), Marram grass (Ammophila breviligulata)
Salt water - green algae, marine eel grass (Zostera spp.), pickleweed (Salicornia virginica), and cordgrass (hybrid Spartina × townsendii) and (Spartina anglica).
Clear water - algae, mosses, freshwater eel grass (Vallisneria americana).
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Ecological succession is the process of change in the species that make up an ecological community over time. The process of succession occurs either after the initial colonization of a newly created habitat, or after a disturbance substantially alters a pre-existing habitat. Succession that begins in new habitats, uninfluenced by pre-existing communities, is called primary succession, whereas succession that follows disruption of a pre-existing community is called secondary succession.
In ecology, habitat refers to the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical manifestation of its ecological niche. Thus "habitat" is a species-specific term, fundamentally different from concepts such as environment or vegetation assemblages, for which the term "habitat-type" is more appropriate. The physical factors may include (for example): soil, moisture, range of temperature, and light intensity.
Primary succession is the beginning step of ecological succession after an extreme disturbance, which usually occurs in an environment devoid of vegetation and other organisms. These environments are typically lacking in soil, as disturbances like lava flow or retreating glaciers scour the environment clear of nutrients. In contrast, secondary succession occurs on substrates that previously supported vegetation before an ecological disturbance.
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