A holotype is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept.
For example, the holotype for the butterfly Plebejus idas longinus is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering.
A holotype is not necessarily "typical" of that taxon, although ideally it is. Sometimes just a fragment of an organism is the holotype, particularly in the case of a fossil. For example, the holotype of Pelorosaurus humerocristatus (Duriatitan), a large herbivorous dinosaur from the early Jurassic period, is a fossil leg bone stored at the Natural History Museum in London. Even if a better specimen is subsequently found, the holotype is not superseded.
Under the ICN, an additional and clarifying type could be designated an epitype under article 9.8, where the original material is demonstrably ambiguous or insufficient.
A conserved type (ICN article 14.3) is sometimes used to correct a problem with a name which has been misapplied; this specimen replaces the original holotype.
In the absence of a holotype, another type may be selected, out of a range of different kinds of type, depending on the case, a lectotype or a neotype.
For example, in both the ICN and the ICZN a neotype is a type that was later appointed in the absence of the original holotype.
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In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen.
In biology, a species (: species) is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour, or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined.
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