Concept

Cupressaceae

Summary
Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecious trees and shrubs up to tall. The bark of mature trees is commonly orange- to red- brown and of stringy texture, often flaking or peeling in vertical strips, but smooth, scaly or hard and square-cracked in some species. The leaves are arranged either spirally, in decussate pairs (opposite pairs, each pair at 90° to the previous pair) or in decussate whorls of three or four, depending on the genus. On young plants, the leaves are needle-like, becoming small and scale-like on mature plants of many genera; some genera and species retain needle-like leaves throughout their lives. Old leaves are mostly not shed individually, but in small sprays of foliage (cladoptosis); exceptions are leaves on the shoots that develop into branches. These leaves eventually fall off individually when the bark starts to flake. Most are evergreen with the leaves persisting 2–10 years, but three genera (Glyptostrobus, Metasequoia and Taxodium) are deciduous or include deciduous species. The seed cones are either woody, leathery, or (in Juniperus) berry-like and fleshy, with one to several ovules per scale. The bract scale and ovuliferous scale are fused together except at the apex, where the bract scale is often visible as a short spine (often called an umbo) on the ovuliferous scale. As with the foliage, the cone scales are arranged spirally, decussate (opposite) or whorled, depending on the genus. The seeds are mostly small and somewhat flattened, with two narrow wings, one down each side of the seed; rarely (e.g. Actinostrobus) triangular in section with three wings; in some genera (e.g. Glyptostrobus and Libocedrus), one of the wings is significantly larger than the other, and in some others (e.g. Juniperus, Microbiota, Platycladus, and Taxodium) the seed is larger and wingless.
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