Concept

Lotka–Volterra equations

Summary
The Lotka–Volterra equations, also known as the Lotka–Volterra predator–prey model, are a pair of first-order nonlinear differential equations, frequently used to describe the dynamics of biological systems in which two species interact, one as a predator and the other as prey. The populations change through time according to the pair of equations: where the variable x is the population density of prey (for example, the number of rabbits per square kilometre); the variable y is the population density of some predator (for example, the number of foxes per square kilometre); and represent the instantaneous growth rates of the two populations; t represents time; The prey's parameters, α and β, describe, respectively, the maximum prey per capita growth rate, and the effect of the presence of predators on the prey growth rate. The predator's parameters, γ, δ, respectively describe the predator's per capita death rate, and the effect of the presence of prey on the predator's growth rate. All parameters are positive and real. The solution of the differential equations is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping. The Lotka–Volterra system of equations is an example of a Kolmogorov model, which is a more general framework that can model the dynamics of ecological systems with predator–prey interactions, competition, disease, and mutualism. The prey are assumed to have an unlimited food supply and to reproduce exponentially, unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation on the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero, then there can be no predation. With these two terms the prey equation above can be interpreted as follows: the rate of change of the prey's population is given by its own growth rate minus the rate at which it is preyed upon.
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