Summary
Pollen is a powdery substance produced by flowers of seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male cone to the female cone of gymnosperms. If pollen lands on a compatible pistil or female cone, it germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail. The study of pollen is called palynology and is highly useful in paleoecology, paleontology, archaeology, and forensics. Pollen in plants is used for transferring haploid male genetic material from the anther of a single flower to the stigma of another in cross-pollination. In a case of self-pollination, this process takes place from the anther of a flower to the stigma of the same flower. Pollen is infrequently used as food and food supplement. Because of agricultural practices, it is often contaminated by agricultural pesticides. Pollen itself is not the male gamete. It is a gametophyte, something that could be considered an entire organism, which then produces the male gamete. Each pollen grain contains vegetative (non-reproductive) cells (only a single cell in most flowering plants but several in other seed plants) and a generative (reproductive) cell. In flowering plants the vegetative tube cell produces the pollen tube, and the generative cell divides to form the two sperm nuclei. Pollen comes in many different shapes. Some pollen grains are based on geodesic polyhedra like a soccer ball. File:Oenothera speciosa pollen 200x.jpg|Triporate pollen of ''[[Oenothera speciosa]]'' File:Lilium auratum - pollen.jpg|Pollen of ''[[Lilium auratum]]'' showing single sulcus (monosulcate) File:Arabis voch1-4.jpg|''[[Arabis]]'' pollen has three colpi and prominent surface structure.
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