Frond

A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the large leaves of cycads, as well as palms (Arecaceae) and various other flowering plants, such as mimosa or sumac. "Frond" is commonly used to identify a large, compound leaf, but if the term is used botanically to refer to the leaves of ferns and algae it may be applied to smaller and undivided leaves. Fronds have particular terms describing their components. Like all leaves, fronds usually have a stalk connecting them to the main stem. In botany, this leaf stalk is generally called a petiole, but in regard to fronds specifically it is called a stipe, and it supports a flattened blade (which may be called a lamina), and the continuation of the stipe into this portion is called the rachis. The blades may be simple (undivided), pinnatifid (deeply incised, but not truly compound), pinnate (compound with the leaflets arranged along a rachis to resemble a feather), or further compound (subdivided). If compound, a frond may be compound once, twice, or more. Pinnation In a frond which is pinnate (feather-shaped), each leafy segment of the blade is called a pinna (plural pinnae), the stalk bearing the pinna is termed a petiolule, and the main vein or mid-rib of the pinna is referred to as a costa (plural costae). If a frond is divided once into pinnae, the frond is called once pinnate. In some fronds the pinna are further divided into segments, creating a bipinnate frond. The segments into which each pinna are divided are called pinnules, and the extensions of the rachis that support these pinnules, are called rachillae. Rarely, a frond may even be tripinnate, in which case the pinnule divisions are known as ultimate segments. Pinnae may be arranged along the rachis either directly opposite one another or alternating up the stem.