Animal echolocation

Echolocation, also called bio sonar, is a biological sonar used by several animal species. Echolocating animals emit calls out to the environment and listen to the echoes of those calls that return from various objects near them. They use these echoes to locate and identify the objects. Echolocation is used for navigation, foraging, and hunting in various environments. Echolocating animals include mammals, most notably Laurasiatheria, especially odontocetes (toothed whales) and some bat species, and also, using simpler forms, species in other groups such as shrews. A few bird species also echolocate, including two cave-dwelling bird groups, the so-called cave swiftlets in the genus Aerodramus (formerly Collocalia) and the unrelated oilbird Steatornis caripensis. The term echolocation was coined by 1944 by the American zoologist Donald Griffin, who, with Robert Galambos, first demonstrated the phenomenon in bats. As Griffin described in his book, the 18th century Italian scientist Lazzaro Spallanzani had, by means of a series of elaborate experiments, concluded that when bats fly at night, they rely on some sense besides vision, but he did not discover that the other sense was hearing. The Swiss physician and naturalist Louis Jurine repeated Spallanzani's experiments (using different species of bat), and concluded that when bats hunt at night, they rely on hearing. In 1908, Walter Louis Hahn confirmed Spallanzani's and Jurine's findings. In 1912, the inventor Hiram Maxim independently proposed that bats used sound below the human auditory range to avoid obstacles. In 1920, the English physiologist Hamilton Hartridge correctly proposed instead that bats used frequencies above the range of human hearing. Echolocation in odontocetes (toothed whales) was not properly described until two decades after Griffin and Galambos' work, by Schevill and McBride in 1956. However, in 1953, Jacques Yves Cousteau suggested in his first book, The Silent World, that porpoises had something like sonar, judging by their navigational abilities.