The Picentes or Piceni or Picentini were an ancient Italic people who lived from the 9th to the 3rd century BC in the area between the Foglia and Aterno rivers, bordered to the west by the Apennines and to the east by the Adriatic coast. Their territory, known as Picenum, therefore included all of today's Marche and the northern part of Abruzzo. The limits of Picenum depend on the era; during the early historic period the region between the Apennines and the Adriatic Sea south of Ancona was Picenum (South Picenians), while between Ancona and Rimini to the north the population was multi-ethnic (North Picenians) because after 390 BC the Senoni Gauls had combined with or supplanted earlier populations. In the Roman Republic the coastal part of northern Picenum was called the ager Gallicus. The origins of the Picentini date from the 9th c. BC as shown by archaeology. They may have been Sabine colonists, although this is doubted by more recent scholars, who see the South Picenes at least as more closely related to the Sabellians. The Piceni did not have a state-type organisation, had no predominant inhabited centre and therefore had no need for a capital. In 390 BC the Senoni Gauls invaded Italy from the north and occupied Picenum north of the Esino river and the centuries-old balance in Picenum underwent drastic changes. The archaeological evidence shows groups of Senones settled much further south of this river, in the Macerata area and even in the Ascoli area, in sites such as Filottrano, San Genesio, Matelica, Offida. When in 299 BC the Romans captured Nequinum, they also concluded a treaty with the Picentes. In 297 BC the Picentes warned the Roman Senate that they had been approached by the Samnites asking for alliance in renewed hostilities with Rome for which the Senate thanked them. The Romans in about 290 BC had absorbed the territory of the Pretuzi, south of Picenum and after a series of victories with the help of the Piceni themselves, the Senones were expelled from the coastal region in 283 BC and the Romans annexed it down to Ancona when it became part of the Ager publicus (Roman state land).