Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing season, and as a means of asexual reproduction. Stem tubers form thickened rhizomes (underground stems) or stolons (horizontal connections between organisms); well known species with stem tubers include the potato and yam. Some writers also treat modified lateral roots (root tubers) under the definition; these are found in sweet potatoes, cassava, and dahlias.
The term originates from the Latin tuber, meaning "lump, bump, swelling".
Some writers define the term "tuber" to mean only structures derived from stems; others use the term for structures derived from stems or roots.
A stem tuber forms from thickened rhizomes or stolons. The top sides of the tuber produce shoots that grow into typical stems and leaves and the undersides produce roots. They tend to form at the sides of the parent plant and are most often located near the soil surface. The underground tuber is normally a short-lived storage and regenerative organ developing from a shoot that branches off a mature plant. The offspring or new tubers are attached to a parent tuber or form at the end of a hypogeogenous (initiated below ground) rhizome. In the autumn the plant dies, except for the new offspring tubers, which have one dominant bud that in spring regrows a new shoot producing stems and leaves; in summer the tubers decay and new tubers begin to grow. Some plants also form smaller tubers and/or tubercules that act like seeds, producing small plants that resemble (in morphology and size) seedlings. Some stem tubers are long-lived, such as those of tuberous begonias, but many plants have tubers that survive only until the plants have fully leafed out, at which point the tuber is reduced to a shriveled-up husk.
Stem tubers generally start off as enlargements of the hypocotyl section of a seedling, but sometimes also include the first node or two of the epicotyl and the upper section of the root.
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In biology, stolons (from Latin stolō, genitive stolōnis – "branch"), also known as runners, are horizontal connections between parts of an organism. They may be part of the organism, or of its skeleton. Typically, animal stolons are exoskeletons (external skeletons). In botany, stolons are plant stems which grow at the soil surface or just below ground that form adventitious roots at the nodes, and new plants from the buds. Stolons are often called runners.
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm or culms. The stem is normally divided into nodes and internodes: The nodes hold one or more leaves, as well as buds which can grow into branches (with leaves, conifer cones, or flowers).
A perennial plant or simply perennial is a plant that lives more than two years. The term (per- + -ennial, "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widely used to distinguish plants with little or no woody growth (secondary growth in girth) from trees and shrubs, which are also technically perennials. Perennialsespecially small flowering plantsthat grow and bloom over the spring and summer, die back every autumn and winter, and then return in the spring from their rootstock or other overwintering structure, are known as herbaceous perennials.
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