Treptichnus (formerly named Phycodes, Manykodes by J. Dzik, and also known as Trichophycus) is the preserved burrow of an animal. As such, it is regarded as the earliest widespread complex trace fossil. Its earliest appearance, around 542 mya, which was contemporaneous with the last of the Ediacaran biota, is used to help define the dividing line, considered geologically at 541 mya, between the Ediacaran and Cambrian periods. It is last seen in the fossil record during the Cenomanian (99.7 Ma).
Treptichnus pedum has a fairly complicated and distinctive burrow pattern: along a central, sometimes sinuous or looping burrow it made successive probes upward through the sediment in search of nutrients, generating a trace pattern reminiscent of a fan or twisted rope. The organism that left such traces is considered more complex than earlier Ediacaran biota; and these trace fossils, which occur worldwide, are usually found in strata above those.
Since only its burrows have been found, it is presumed that the Treptichnus animal lacked any hard anatomical features, such as shells or bones. Its morphology and relationship to modern animals is therefore unknown, and some dispute even its inclusion within the animal kingdom. Despite this lack of fossil evidence, surface trails produced by modern priapulid worms exhibit the same probing morphology as Treptichnus, suggesting a close anatomical relationship between the trace-maker and modern priapulids.
The name Treptichnus pedum means "turned-trail (Greek) of feet (Latin)".
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The Ediacaran (ˌiːdiˈækərən; formerly Vendian) biota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period ( 635–538.8 Mya). These were enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.
The Cryogenian (from krýos, meaning "cold" and γένεσις, romanized: , meaning "birth") is a geologic period that lasted from . It forms the second geologic period of the Neoproterozoic Era, preceded by the Tonian Period and followed by the Ediacaran. The Cryogenian was a time of drastic biosphere changes. After the previous Boring Billion years of stability, at the beginning of Cryogenian the severe Sturtian glaciation began, freezing the entire Earth in a planetary state known as a Snowball Earth.
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