The Ati are a Negrito ethnic group in the Visayas, the central portion of the Philippine archipelago. Their small numbers are principally concentrated in the islands of Boracay, Panay and Negros. They are genetically related to other Negrito ethnic groups in the Philippines such as the Aeta of Luzon, the Batak of Palawan, the Agta of the Sierra Madres, and the Mamanwa of Mindanao. The Negritos are the descendants of the same early "East-Eurasian" meta-population, which also gave rise to modern East Asians and Australasians, among other populations of the Asia-Pacific region. The earliest modern human migrations into the Philippine archipelago were during the Paleolithic, around 40,000 years ago, followed by two other migration waves between 25,000 and 12,000 years ago, through the Sundaland land bridges that linked the islands with the Asian mainland. The latest migration wave is associated with the Austronesian peoples (c. 7,000 years ago) from Taiwan. The Philippine Negritos display relatively closer genetic affinity towards different Eastern Asian populations, prehistoric Hoabinhian samples, as well as to the Indigenous people of New Guinea and Aboriginal Australians, from which they diverged around c. 40,000 years ago, and also display genetic substructure along a North to South cline, suggesting their ancestral population diverged into two subgroups after the initial peopling of the Philippines. Furthermore, they display high percentages of Denisovan gene flow. Legends, such as those involving the Ten Bornean Datus and the Binirayan Festival, tell tales about how, at the beginning of the 12th century when Indonesia and the Philippines were under the rule of Indianized native kingdoms, the ancestors of the Bisaya escaped from Borneo and from the persecution of Rajah Makatunaw. Led by Datu Puti and Datu Sumakwel and sailing with boats called balangays, they landed near a river called Suaragan, on the southwest coast of Panay, (the place then known as Aninipay), and bartered the land from an Ati headman named Polpolan and his son Marikudo for the price of a necklace and one golden salakot.