Concept

Vernalization

Summary
Vernalization () is the induction of a plant's flowering process by exposure to the prolonged cold of winter, or by an artificial equivalent. After vernalization, plants have acquired the ability to flower, but they may require additional seasonal cues or weeks of growth before they will actually do so. The term is sometimes used to refer to the need of herbal (non-woody) plants for a period of cold dormancy in order to produce new shoots and leaves, but this usage is discouraged. Many plants grown in temperate climates require vernalization and must experience a period of low winter temperature to initiate or accelerate the flowering process. This ensures that reproductive development and seed production occurs in spring and winters, rather than in autumn. The needed cold is often expressed in chill hours. Typical vernalization temperatures are between 1 and 7 degrees Celsius (34 and 45 degrees Fahrenheit). For many perennial plants, such as fruit tree species, a period of cold is needed first to induce dormancy and then later, after the requisite period of time, re-emerge from that dormancy prior to flowering. Many monocarpic winter annuals and biennials, including some ecotypes of Arabidopsis thaliana and winter cereals such as wheat, must go through a prolonged period of cold before flowering occurs. In the history of agriculture, farmers observed a traditional distinction between "winter cereals", whose seeds require chilling (to trigger their subsequent emergence and growth), and "spring cereals", whose seeds can be sown in spring, and germinate, and then flower soon thereafter. Scientists in the early 19th century had discussed how some plants needed cold temperatures to flower. In 1857 an American agriculturist John Hancock Klippart, Secretary of the Ohio Board of Agriculture, reported the importance and effect of winter temperature on the germination of wheat. One of the most significant works was by a German plant physiologist Gustav Gassner who made a detailed discussion in his 1918 paper.
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