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Concept
Vacuole
Summary
A vacuole (ˈvækjuːoʊl) is a membrane-bound organelle which is present in plant and fungal cells and some protist, animal, and bacterial cells. Vacuoles are essentially enclosed compartments which are filled with water containing inorganic and organic molecules including enzymes in solution, though in certain cases they may contain solids which have been engulfed. Vacuoles are formed by the fusion of multiple membrane vesicles and are effectively just larger forms of these. The organelle has no basic shape or size; its structure varies according to the requirements of the cell.
Contractile vacuoles ("stars") were first observed by Spallanzani (1776) in protozoa, although mistaken for respiratory organs. Dujardin (1841) named these "stars" as vacuoles. In 1842, Schleiden applied the term for plant cells, to distinguish the structure with cell sap from the rest of the protoplasm.
In 1885, de Vries named the vacuole membrane as tonoplast.
The function and significance of vacuoles varies greatly according to the type of cell in which they are present, having much greater prominence in the cells of plants, fungi and certain protists than those of animals and bacteria. In general, the functions of the vacuole include:
Isolating materials that might be harmful or a threat to the cell
Containing waste products
Containing water in plant cells
Maintaining internal hydrostatic pressure or turgor within the cell
Maintaining an acidic internal pH
Containing small molecules
Exporting unwanted substances from the cell
Allows plants to support structures such as leaves and flowers due to the pressure of the central vacuole
By increasing in size, allows the germinating plant or its organs (such as leaves) to grow very quickly and using up mostly just water.
In seeds, stored proteins needed for germination are kept in 'protein bodies', which are modified vacuoles.
Vacuoles also play a major role in autophagy, maintaining a balance between biogenesis (production) and degradation (or turnover), of many substances and cell structures in certain organisms.
Official source
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