Concept

Evolution of the eye

Many scientists have found the evolution of the eye attractive to study because the eye distinctively exemplifies an analogous organ found in many animal forms. Simple light detection is found in bacteria, single-celled organisms, plants and animals. Complex, image-forming eyes have evolved independently several times. Diverse eyes are known from the Burgess shale of the Middle Cambrian, and from the slightly older Emu Bay Shale. Eyes vary in their visual acuity, the range of wavelengths they can detect, their sensitivity in low light, their ability to detect motion or to resolve objects, and whether they can discriminate colours. In 1802, philosopher William Paley called it a miracle of "design." In 1859, Charles Darwin himself wrote in his Origin of Species, that the evolution of the eye by natural selection seemed at first glance "absurd in the highest possible degree". However, he went on that despite the difficulty in imagining it, its evolution was perfectly feasible: if numerous gradations from a simple and imperfect eye to one complex and perfect can be shown to exist, each grade being useful to its possessor, as is certainly the case; if further, the eye ever varies and the variations be inherited, as is likewise certainly the case and if such variations should be useful to any animal under changing conditions of life, then the difficulty of believing that a perfect and complex eye could be formed by natural selection, though insuperable by our imagination, should not be considered as subversive of the theory. He suggested a stepwise evolution from "an optic nerve merely coated with pigment, and without any other mechanism" to "a moderately high stage of perfection", and gave examples of existing intermediate steps. Current research is investigating the genetic mechanisms underlying eye development and evolution. Biologist D.E. Nilsson has independently theorized about four general stages in the evolution of a vertebrate eye from a patch of photoreceptors. Nilsson and S.

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Related concepts (5)
Cambrian explosion
The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period of early Paleozoic when there was a sudden radiation of complex life and practically all major animal phyla started appearing in the fossil record. It lasted for about 13 – 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.
Arthropod
Arthropods (ˈɑːrθrəpɒd, (gen. ποδός)) are invertebrate animals in the phylum Arthropoda. They possess an exoskeleton with a cuticle made of chitin, often mineralised with calcium carbonate, a segmented body, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They are an extremely diverse group, with up to 10 million species. Haemolymph is the analogue of blood for arthropods.
Simple eye in invertebrates
A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-lensed "compound eye", and is not necessarily at all simple in the usual sense of the word. The structure of an animal's eye is determined by the environment in which it lives, and the behavioural tasks it must fulfill to survive.
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