In a vascular plant, the stele is the central part of the root or stem containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue (pith) and a pericycle, which, if present, defines the outermost boundary of the stele. Outside the stele lies the endodermis, which is the innermost cell layer of the cortex.
The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. Now, at the beginning of the 21st century, plant molecular biologists are coming to understand the genetics and developmental pathways that govern tissue patterns in the stele. Moreover, physiologists are examining how the anatomy (sizes and shapes) of different steles affect the function of organs.
The earliest vascular plants had stems with a central core of vascular tissue. This consisted of a cylindrical strand of xylem, surrounded by a region of phloem. Around the vascular tissue there might have been an endodermis that regulated the flow of water into and out of the vascular system. Such an arrangement is termed a protostele.
There are usually three basic types of protostele:
haplostele – consisting of a cylindrical core of xylem surrounded by a ring of phloem. An endodermis generally surrounds the stele. A centrarch (protoxylem in the center of a metaxylem cylinder) haplostele is prevalent in members of the rhyniophyte grade, such as Rhynia.
actinostele – a variation of the protostele in which the core is lobed or fluted. This stele is found in many species of club moss (Lycopodium and related genera). Actinosteles are typically exarch (protoxylem external to the metaxylem) and consist of several to many patches of protoxylem at the tips of the lobes of the metaxylem. Exarch protosteles are a defining characteristic of the lycophyte lineage.
This page is automatically generated and may contain information that is not correct, complete, up-to-date, or relevant to your search query. The same applies to every other page on this website. Please make sure to verify the information with EPFL's official sources.
The presentation of tree growth and formation of wood anatomical structures, linked to the description of specific physical and mechanical properties, makes it possible to understand the different for
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm or culms. The stem is normally divided into nodes and internodes: The nodes hold one or more leaves, as well as buds which can grow into branches (with leaves, conifer cones, or flowers).
Aglaophyton major (or more correctly Aglaophyton majus) was the sporophyte generation of a diplohaplontic, pre-vascular, axial, free-sporing land plant of the Lower Devonian (Pragian stage, around ). It had anatomical features intermediate between those of the bryophytes and vascular plants or tracheophytes. A. major was first described by Kidston and Lang in 1920 as the new species Rhynia major.
The rhyniophytes are a group of extinct early vascular plants that are considered to be similar to the genus Rhynia, found in the Early Devonian (around ). Sources vary in the name and rank used for this group, some treating it as the class Rhyniopsida, others as the subdivision Rhyniophytina or the division Rhyniophyta. The first definition of the group, under the name Rhyniophytina, was by Banks, since when there have been many redefinitions, including by Banks himself.
Alfalfa leaf curl virus (ALCV) is the first geminivirus for which aphid transmission was reported. Transmission by Aphis craccivora was determined previously to be highly specific and circulative. Using various complementary techniques, the transmission jo ...
Plants close their stomata during drought to avoid excessive water loss, but species differ in respect to the drought severity at which stomata close. The stomatal closure point is related to xylem anatomy and vulnerability to embolism, but it also has imp ...
2018
,
The stable isotopes of hydrogen and oxygen in xylem water are often used to investigate tree water sources. But this traditional approach does not acknowledge the contribution of water stored in the phloem to transpiration and how this may affect xylem wat ...